After completing Light the Dragonfires, it is possible to obtain a second blessing from the statue by activating it from outside the Temple (a high Acrobatics skill is necessary). This blessing fortifies Personality and Luck each for 86,400 seconds (one full real-time day). After completing the Main Quest, you can receive the blessing Favor of Akatosh from the statue of the Avatar. During the quest "Light the Dragonfires", the Staff of Sheogorath will be able to affect the entire temple. Another possible source is the dome of St Peter's Basilica, designed by Michelangelo, which has a similar arrangement of projecting paired columns. The Temple of the One is similar to the Pantheon of Rome and might be based on it, judging by the fact that Imperials are based on Romans. A Vampire in later stages will take sun damage inside the Temple of the One. If the Hero saved Jeelius from the Mythic Dawn sacrificial ceremony inside the Lake Arrius Caverns, he appears at the Temple. Sometimes, for unknown reasons, after using the statue of the avatar of Akatosh and receiving the blessing, the timer will not go down and the effects will be permanent. It is a glitchy way to finish the main quest quickly. 
If one walks outside, they will see the Temple District overrun by Daedra. If one goes inside, they will see the temple as it is during the end of the main quest. There is a half-buried door right behind where the normal door is.
Before one finishes the main quest, if their Acrobatics skill is high enough, one may be able to jump onto the temple, fall through the roof and into the empty area inside the walls. If one succeeds, they will drop down until they spawn back in the center of the temple. After completing the main quest, with a high enough Acrobatics, one can jump on one of the pillars, then on the leg of the dragon, and across the wall. The Altmer, Tandilwe, requests the Hero to talk to every beggar in the Imperial City, before she can offer training in Speechcraft. With the Amulet of Kings in hand, it is time for Martin Septim to return to the Imperial City, claim the Septim Throne, and light the Dragonfires in order to stop the Daedric invasion. 
J'mhad, the groundskeeper, and Tandilwe, the master speechcraft trainer, as well as several other worshippers can be found inside the temple throughout the day. During the Oblivion Crisis, the Temple was the site of the final showdown between Mehrunes Dagon and the avatar of Akatosh, which leaves the roof of the temple shattered, debris and damage scattered throughout the surrounding streets, and the avatar of Akatosh encased in stone.
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